Riccardo Papa, University of Puerto Rico (IMAGE) U.S. National Science Foundation Caption Riccardo Papa, of the University of Puerto Rico Rio Piedras, in the field. The university's project uses butterfly wing color patterns as a mechanism to study genotype-phenotype relationships in Heliconius butterflies. Credit Riccardo Papa, University of Puerto Rico, Department of Biology. Usage Restrictions None License Licensed content Disclaimer: AAAS and EurekAlert! are not responsible for the accuracy of news releases posted to EurekAlert! by contributing institutions or for the use of any information through the EurekAlert system.