Ruffed grouse illustration (IMAGE) Penn State Caption Pennsylvania’s ruffed grouse populations have declined by up to 70% since the early 1960s, with birds in the southern part of the state particularly affected by West Nile virus, which is spread by mosquitoes, and by habitat fragmentation due to development. Credit Andreza Silva Usage Restrictions Educational use only License Original content Disclaimer: AAAS and EurekAlert! are not responsible for the accuracy of news releases posted to EurekAlert! by contributing institutions or for the use of any information through the EurekAlert system.