Riccardo Papa, University of Puerto Rico (image) National Science Foundation Share Print E-Mail Caption Riccardo Papa, of the University of Puerto Rico Rio Piedras, in the field. The university's project uses butterfly wing color patterns as a mechanism to study genotype-phenotype relationships in Heliconius butterflies. Credit Riccardo Papa, University of Puerto Rico, Department of Biology. Usage Restrictions None Share Print E-Mail Disclaimer: AAAS and EurekAlert! are not responsible for the accuracy of news releases posted to EurekAlert! by contributing institutions or for the use of any information through the EurekAlert system.