Subordinate Hyena Giggle Call (AUDIO) University of California - Berkeley Your browser does not support the audio element. Caption The giggle call of a subordinate male hyena named Winnie (who was 14 years old at the time of the study) is more variable and tentative than the steady call of dominant animals. Credit Frederic Theunissen/UC Berkeley Usage Restrictions None License Licensed content Disclaimer: AAAS and EurekAlert! are not responsible for the accuracy of news releases posted to EurekAlert! by contributing institutions or for the use of any information through the EurekAlert system.