Cladogram of the Tribes (IMAGE) Pensoft Publishers Caption Cladogram of the tribes Oniticellini (left) and Onthophagini (right). Credit T. Keith Philips Usage Restrictions CC-BY 4.0 License Licensed content Disclaimer: AAAS and EurekAlert! are not responsible for the accuracy of news releases posted to EurekAlert! by contributing institutions or for the use of any information through the EurekAlert system.